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Species traits (environmental optima, niche breadths and\nmaximum growth rates) can carry phylogenetic signal or evolve\nunder an Ornstein-Uhlenbeck process with one or several optima,\nand interspecific competition can be structured by relatedness.\nRule-based assembly without population dynamics (limiting\nsimilarity, habitat filtering and neutral assembly) follows\nKraft et al. (2007) , and static\nmetacommunities with unimodal species responses follow\nPeres-Neto et al. (2012) . 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